Total dry matter production
Intercropping with pulses had significant effect on the total dry matter production (TDMP) of maize and finger millet, intercropped in coconut garden (Table 1).
Table 1: TDMP and yield (pooled mean) of maize, finger millet, black gram and green gram as influenced by intercropping in coconut garden, kg ha-1.
Between the sole crops, maize recorded higher total dry matter production (12626 kg ha-1
) than finger millet (4942 kg ha-1
). The robust nature of maize as evidenced by taller plants and larger leaves, compared to finger millet might have contributed to the higher dry matter production. Intercropping maize with black gram resulted in 16.91 per cent higher TDMP of maize (11234 kg ha-1
), than intercropping with green gram (10747 kg ha-1
). Intercropping with black gram was observed to increase the TDMP of finger millet (3984 kg ha-1
) by 5.17 per cent than green gram. (3788 kg ha-1
Sole crops of black gram and green gram recorded significantly higher TDMP (1916 kg ha-1
and 2300 kg ha-1
) compared to intercropping. In general, cereals and millets possess greater rooting densities (Anil et al., 1998).
Thus, when pulses were intercropped with maize and finger millet, a competition might have emerged affecting the growth of pulses when intercropped, as reported by Reddy et al., (2021).
Both black gram and green gram resulted in statistically comparable TDMP, when intercropped. However, the TDMP of green gram was observed to be higher (1441 kg ha-1
) than black gram (1204 kg ha-1
) under intercropping.
Sole crops of maize and finger millet intercropped in coconut garden recorded significantly higher grain yields (6880 kg ha-1
and 1760 kg ha-1 respectively) (Table 1), than when intercropped with pulses (5177 kg ha-1
and 1372 kg ha-1
). While difference between the sole crop and intercrop yields was 32.9 per cent for maize, it was 28.3 per cent for finger millet. Plant population is a key factor that decides yield. Thus it is logical to reason out that the higher plant population of maize and finger millet under sole cropping might have contributed to the higher yields. The reduction in the yields under intercropping could also be attributed to the competition with the pulses along with a reduced plant population. Maize + black gram resulted in 7.10 per cent higher grain yield of maize (5354 kg ha-1
) than maize + green gram, (4999 kg ha-1
). The yield of finger millet was statistically comparable between the two intercrops, viz
., black gram and green gram.
There were significant variations in the haulm yields of maize and finger millet intercropped in coconut garden (Table 1). Haulm yield was also significantly superior for the sole crops of maize and finger millet (9211 kg ha-1
and 4053 kg ha-1
). Haulm yield of both maize and finger millet were superior when intercropped with black gram (8898 kg ha-1
and 3280 kg ha-1
Intercropping had significant effect on the seed yield of black gram and green gram (Table 1), with higher yields under sole cropping. Sole crop yields of black gram and green gram were almost equal. However, under intercropping, black gram yielded higher than green gram. The seed yield of black gram under finger millet + black gram (501 kg ha-1
) was comparable with maize + black gram (495 kg ha-1
). Green gram also behaved in a similar manner with higher seed yield under finger millet + green gram (457 kg ha-1
) followed by maize + green gram (357 kg ha-1
). The higher seed yield of black gram might be due to the higher test weight of black gram compared to green gram. Similar observations have been made by Sarma et al., (2016)
in studies conducted with intercropping black gram and green gram in sesame.
The bhusa yields were also superior for the sole crops of black gram (1499 kg ha-1
) and green gram (2496 kg ha-1
). Bhusa yield of black gram was statistically comparable when intercropped with finger millet (1005 kg ha-1
) and maize (988 kg ha-1
). Bhusa yield of green gram was also comparable when intercropped with finger millet (1736 kg ha-1
) and maize (1623 kg ha-1
Under intercropping, the mean seed yield of black gram was 498 kg ha-1
and that of green gram was 407 kg ha-1
. The seed yield of black gram was 22.4 per cent higher than green gram under intercropping. Although the TDMP of green gram was 19.7 per cent higher than black gram, this was not reflected in the seed yield of the green gram. This is possibly because of a better source to sink translocation in black gram than in green gram.
Land equivalent ratio
The land equivalent ratio (LER) indicates the advantage of an intercropping system with efficient resource utilization compared to pure stands of the respective crops. LER values greater than unity denotes yield advantage. The data on the effect of intercropping on LER are presented in Table 2.
Table 2: Effect of intercropping on land equivalent ratio (LER), aggressivity (A) and percentage yield difference (PYD).
The LER was higher for maize + black gram, closely followed by finger millet + black gram. The inclusion of black gram as an intercrop in maize or finger millet resulted in on an average 35 per cent yield advantage. According to Vandermeer (1989)
, intercrops that result in LER values greater than unity are considered to over yield, gaining their advantage through the ‘competitive production principle’ and/or the ‘facilitative production principle’. The higher LER in intercropping than sole cropping could be attributed to the better utilization of both natural and supplemented resources.
Aggressivity (A) is a focal competitive function used to assess the competitive ability of a crop when grown along with another crop. Aggressivity values with positive (+) sign denotes the dominant species and negative (-) sign denotes the dominated species. When the numerical values are greater, it indicates greater difference in the competitive abilities of the component crops. The data presented in Table 2 showed that the competitive ability of the component crops varied widely. Maize was more competitive than finger millet against pulses, as indicated by the positive (+) sign. The C4 nature of maize might have contributed to its competitive behavior (Kheroar and Patra, 2014)
The competitive ability of maize and black gram was observed to be closer as indicated by lower values than maize + green gram. Maize tended to be more competitive than green gram. On the contrary, pulses exhibited more competitiveness with finger millet. Black gram showed greater competition with finger millet than green gram.
Percentage yield difference
Percentage yield difference (PYD) is an index of the yield compensation capacity of the component crops in an intercropping system. Irrespective of the proportion of crops, planting time and planting geometry, the reduction in yield of one crop gets compensated with increase in yield of the other. The PYD was the highest (36.36%) for finger millet + black gram, followed by maize + black gram (33.56%) (Table 2). This pointed towards the fact that the yield reduction of maize and finger millet under intercropping was compensated by increase in yield of black gram in a better manner than green gram. The superiority of black gram as intercrop with maize and finger millet has been reported previously by Kheroar and Patra (2014) and Kumar and Ray (2020)
Coconut equivalent yield
Crop equivalent yield has been identified as one among the efficient indices capable of assessing the overall production potential of intercropping systems. Coconut equivalent yield (CEY) was higher for intercropping than sole cropping. Among the coconut based intercropping systems, CEY was significantly the highest (5500 kg ha-1
) for intercropping coconut with maize + black gram, followed by finger millet + black gram (5289 kg ha-1
) (Table 3).
Table 3: Effect of intercropping on coconut equivalent yield, kg ha-1.
With respect to intercropping with green gram, the coconut equivalent yield was higher (4823 kg ha-1
) when green gram was intercropped with finger millet, than with maize (4780 kg ha-1
). When the sole crops were compared, it was observed that the production potential of coconut garden (2250 kg ha-1
) could be increased significantly when black gram was intercropped in coconut garden as indicated by significantly higher CEY (4787 kg ha-1
), followed by maize (4609 kg ha-1
Nitrogen balance sheet
The balance sheet of available nitrogen (N) was observed to be negative following the sole crops of maize and finger millet and maize + pulses (Fig 1).
Fig 1: Balance sheet of available N, P and K as influenced by intercropping, kg ha-1.
C- Coconut; M- Maize; FM- Finger millet; BG- Black gram; GG- Green gram.
The depletion of N was observed to be more for maize. The depletion in N following maize could be attributed to the higher uptake of N due to its exhaustive nature. However, when maize was intercropped with pulses, there was a lessening in the depletion of N as indicated by lesser negative values. Several studies have shown that maize is a crop that places major demand for resources at different times of system duration. When legumes or pulses are intercropped with maize, the longer duration maize could recover its resource needs after the harvest of the pulses. A positive balance was observed for available N after the sole crops of black gram and green gram. Finger millet + pulses also resulted in positive balance for available N in soil. Legumes derive 5 per cent to 83 per cent of their N requirement by fixing atmospheric nitrogen under field conditions and remaining from the soil N pool (Dudeja and Duhan, 2005)
. Thus the demand placed by legumes on soil N is less compared to the non-nitrogen fixing crops like maize and finger millet. This together with the legume effect might have resulted in the positive balance for available N in the soil. Between the two pulses, green gram left the soil more enriched in terms of N, both under sole crop and as intercrop.
Phosphorus balance sheet
The data on the balance sheet of available phosphorus (P) are presented in Fig 1. Balance sheet of available (P) was observed to be negative in all the treatments, including sole crops and intercrops. The magnitude of depletion was higher for sole crop of maize and in intercropping systems involving maize. Studies have shown rhizosphere acidification and increase of DTPA - extractable Fe and concomitant decrease in rhizosphere phosphorus in maize. Further, the soil of the experimental site was acidic in reaction. The negative balance of P observed might be due to P fixation under acidic pH. Similar cases P fixation as iron and aluminium phosphate was reported by Huck et al., (2014).
Potassium balance sheet
The balance sheet of available potassium (K) was observed to be positive in all the treatments, except black gram and green gram raised as sole crops in coconut garden (Fig 1). The initial status of available K was rated as low. Potassium may be lost from soil through leaching. Both these factors might have contributed to a negative balance for available K after pulses. In the case of maize and finger millet and the respective intercropping systems with pulses, the computed balance was lower than the actual balance, leading to net gain. Maize and finger millet, being members of the Poaceae family require larger quantity of K for their growth and development. Thus their higher values of K uptake led to lower computed balance for potassium than its actual balance.
On the whole, the nutrient balance sheet revealed the importance of soil test based nutrient management in coconut garden. Though maize proved to be an exhaustive crop as hypothesized, intercropping maize with pulses was observed to compensate for the exhaustive behavior of maize. Pulses were observed to improve the fertility status of the soil.