Growth characteristics
The differences in plant height and dry matter accumulation of blackgram due to different intercropping systems were found to be significant at 60 DAS (Table 2). The higher plant height (45.80 cm) and dry matter accumulation (7.03 g plant
-1) were recorded under sole blackgram, which was statistically at par with blackgram + sesame 5:1 replacement series, blackgram + sesame 3:1 replacement series, blackgram + sesame 1:1 replacement series and remain significantly superior over all other treatments. However, the lowest plant height and dry matter accumulation of blackgram were recorded in blackgram + sesame additive series. Significant changes in plant height and dry matter accumulation might have occurred due to competitiveness of plants for various essentials. The reduced competition for light, nutrients and other essentials under sole blackgram which probably might have led to increased height.
Chhetri et al., (2015) also reported that there was significant reduction in plant height of blackgram when they were grown with sesame in an intercropping system as compared to their sole treatment.
The different intercropping systems showed significant difference in plant height and dry matter accumulation of sesame at 60 DAS (Table 2). Among the different intercropping systems highest plant height and dry matter accumulation to the tune of 107.73 cm 18.93 g plant
-1, respectively were observed in blackgram + sesame 5:1 replacement series, which was statistically at par with blackgram + sesame 3:1 replacement series, sole sesame, blackgram + sesame 1:1 replacement series, blackgram + sesame 1:3 replacement series and blackgram + sesame 1:5 replacement series of intercropping system but significantly superior over blackgram + sesame seed mix series and blackgram + sesame additive series. However, the lowest plant height (93.65 cm) and dry matter accumulation (11.09 g plant
-1) were recorded with blackgram + sesame additive series. It was probably due to more space and nutrients available for growth and development of sesame, which led to higher photosynthesis owing to greater exposure of sesame plants to sunlight. It might also be due to conducive environment created by main crop (blackgram) as it fixed atmospheric nitrogen and increased its availability in soil which might have also been utilized partly by sesame plants for better growth and development and ultimately increased the growth characteristics. Similar results were also reported by
Meena et al., (2008) in intercropping of sesame and cluster bean.
Yield attributes and yield
The intercropping of sesame had significant effect on yield attributes (pods plant
-1 and seeds pod
-1) of blackgram. A comparison of yield attributing characters of the different blackgram based intercropping systems showed that the sole crop of blackgram recorded highest plants m
-2 (30.6) followed by additive series treatment (Table 3). Higher number of pods plant
-1, seeds pod
-1 and 1000-seed weight of sole stand of blackgram being statistically similar with blackgram + sesame replacement series of 5:1, 3:1, 1:1 and 1:3 row ratio but superior to blackgram + sesame 1:5 replacement series, blackgram + sesame additive series and seed mix series of blackgram + sesame intercropping. Similar results of increasing yield attributes were also reported by
Sarkar et al., (2000). The data given in Table 4 revealed that blackgram in sole recorded highest seed and stover yield (5.88 q ha
-1) and (18.81 q ha
-1) and was followed by blackgram + sesame replacement series of 5:1 row ratio. Whereas, lowest seed and stover yield was obtained in blackgram + sesame 1:5 replacement series. The higher seed and stover yield of blackgram sole and 1:5 replacement series might have happened due to higher plant population as compared to other treatments. The reduction in seed and stover yields of blackgram in all intercropping systems over sole blackgram was primarily due to low plant population of blackgram in intercropping treatments. Reduction in yield of main crop blackgram might also be due to shading effect of sesame (component crop) on blackgram. These results are in close conformity with those of
Tripathi et al., (2005) who reported significant reduction in yield of chickpea when grown in association with mustard. The blackgram seed yield increases with increase in pods plant
-1 and 1000 seed weight of blackgram (Fig 1a and b).
A comparison of yield attributing characters of different intercropping treatments showed that the blackgram + sesame in 5:1 row ratio recorded highest capsule plant
-1, seeds capsule
-1 and 1000-seed weight of sesame which was at par with blackgram + sesame 3:1 replacement series, blackgram + sesame 1:1 replacement series, blackgram + sesame 1:3 replacement series and blackgram + sesame 1:5 replacement series (Table 3). It was probably due to more space and nutrients available for growth and development of sesame, which led to higher photosynthesis owing to greater exposure of sesame plants to sunlight and also be due to conducive environment created by main crop (blackgram) as it fixed atmospheric nitrogen and increased its availability in soil which might have also been utilized partly by sesame plants for better growth, development and yield attributes. Similar results were also reported by
Meena et al., (2008) in intercropping of blackgram and sesame. The data presented in Table 4 revealed that among the different intercropping systems, the sole planting of sesame gave significantly higher seed (3.47 q ha
-1) and stick yields (15.84 q ha
-1) followed by blackgram + sesame in 1:5 series as compared to other planting in different row ratios. This was primarily due to higher plant population per unit area in sole sesame. The higher yields under pure stand were also reported by
Tiwari et al., (1994) in sesame, greengram and soybean and Sarkar and Sanyal (2000) in sesame, groundnut and sunflower as compared to their yields under intercropping systems. The sesame seed yield increases with increase in capsules plant
-1 and 1000 seed weight of sesame (Fig 1c and d).
In blackgram and sesame intercropping system, sesame yield decreases with increase in blackgram yield (Fig 1e). Blackgram equivalent yield (BEY) significantly influenced by different intercropping systems (Table 4). However, among the different intercropping systems, blackgram + sesame 5:1 replacement series recorded significantly higher blackgram equivalent yield which was at par with blackgram + sesame 3:1 replacement series, blackgram + sesame additive series and blackgram + sesame 1:1 replacement series. The lowest blackgram equivalent yield was calculated in seed mix series blackgram + sesame intercropping system. This increase in blackgram equivalent yield of intercropping systems over sole blackgram system was mainly due to beneficial effects of intercropping with differential yield behaviours of the crops which ensured higher total productivity and profitability due to additional yield advantage of intercrop yield.
Prajapat et al., (2012) also found higher values of mungbean equivalent yield in mungbean+sesame intercropping as compared to sole sesame.
Tripathi et al., (2005) and
Ahlawat et al., (2005) also recorded similar findings in respect of chickpea equivalent yield in chickpea+mustard intercropping system.
Intercropping indices
All the intercropping treatments resulted in significantly higher LER as compared to the sole crop (Table 5). Blackgram + sesame in 5:1 replacement series gave the highest LER upto (1.22) followed by blackgram + sesame 3:1 replacement series (1.14) and blackgram + sesame additive series (1.11) and found biologically more efficient as compared to other intercropping systems. This yield advantage due to intercropping could possibly be attributed to the combined effects of better utilization of resources by component crops having different rooting patterns, differential canopy distribution and efficient light interception by their green surfaces and differential nutrient extraction from different soil depths in intercropping system. Whereas, lowest value 0.78 of LER was realized in blackgram + sesame (seed mix). The area-time equivalent ratio (ATER) is the performance of each intercrop component by the length of time required to grow and harvest it. Blackgram + sesame in 5:1 replacement series recorded higher value of ATER (1.14) as compared to other replacement and additive treatment (Table 5). This might have happened due to optimum utilization of land resources with respect to time 5:1 replacement series treatment. These results were in accordance with the findings of
Sinha et al., (1999) and
Tripathi et al., (2010).The competitive ability of the component crops in an intercropping system is determined by its aggressivity value. The zero value of aggressivity indicates that component crops are equally competitive. For any other situation, both crops will have the same numerical value but the sign of the dominant species will be positive and that of dominated negative. Among the intercropping treatments, positive value of 0.66, 0.46, 0.26, 0.17 0.13 and 0.02 were recorded in sesame crop of replacement series of 5:1, 1:5, 3:1, 1:3, seed mix and 1:1 blackgram + sesame intercropping systems, respectively, which denotes in all these treatments sesame crop was dominant on blackgram (Table 5). This probably happened due to early suppressive ability of the fast growing high foliage sesame crop along with its better ability to intercept light and also utilize soil resources which enabled it to become more efficient in resource utilization as compared to blackgram crop. Negative value of -0.29 was recorded in crop in additive treatment which showed that in this case blackgram crop was dominant on sesame crop. Among the intercropping treatments, blackgram + sesame 5:1 replacement series registered highest net returns and B: C ratio followed by blackgram + sesame 3:1 replacement series. However, the seed mix series of blackgram + sesame intercropping system registered lowest net returns (Table 4).
Soil fertility changes
The soil chemical parameters
viz. pH, EC and organic carbon and availability of nutrients (N, P and K) in soil after harvesting of blackgram and component crop (sesame) were not influenced significantly by different intercropping systems (Table 6). However, values of available N, P and K in soil parameters were observed to be higher in additive series treatment followed by sole blackgram and different replacement series and in sole sesame. Similar findings were also reported by
Khola et al., (2000).
Energy
Energy in agriculture is important in terms of crop production and agro processing for value adding (Karimi
et. al., 2008). The relation between agriculture and energy is very close. Agriculture itself is an energy user and energy supplier in the form of bio-energy. At present, productivity and profitability of agriculture depends on energy consumption
(Alam et al., 2005). Data presented in Table 7 revealed that among the different treatments energy output (MJ ha
-1), energy use efficiency and energy intensity (MJ Rs.
-1) were significantly higher with blackgram+sesame 5:1(replacement series) which was statistically at par with blackgram + sesame 3:1(replacement series) and blackgram + sesame additive series. The net energy return (MJ ha
-1) was significantly higher with blackgram + sesame 5:1 (replacement series) than all other treatments. The energy productivity (kg MJ
-1) was significantly higher with blackgram + sesame 5:1 (replacement series) which was statistically at par with blackgram + sesame 3:1 (replacement series), blackgram + sesame 1:1 (replacement series), blackgram + sesame additive series and sole sesame and blackgram.The higher blackgram equivalent yield under blackgram + sesame 5:1 (replacement series) and blackgram + sesame 3:1 (replacement series) resulted in higher energy output, energy use efficiency, energy productivity, net energy return and energy intensity. However, the specific energy (MJ kg
-1) was found significantly highest with blackgram + sesame (seed mix) than all other treatments.