Effect of intercropping on growth attributes
The data on growth attributes of groundnut are presented in Table 1. An appraisal of data revealed that growth attributes of groundnut were significantly influenced by mix/intercropping system in pooled results of both the years 2020-21 and 2021-22.
Among the different treatments, sole groundnut (T1) had recorded significantly the highest growth attributes of groundnut
viz., final plant population (22.26 plants/m
2), number of branches/plant (7.97), dry matter production/plant (37.37 g) and number (80.9) and fresh weight (3.75 g) of root nodules/plant. Whereas, significantly highest plant height (67.84 cm) was recorded with intercropping systems of groundnut + sweet corn (1:1) in additive series (T6) and maximum leaf area/plant (2911.4 cm
2) and leaf area index (6.470) of groundnut at harvest was recorded in paired row sowing of groundnut + sweet corn (2:1) at 30-60-30 cm (T10) in pooled results. Intercropping treatments did not show their significantly effected on plant spread.
In additive intercropping systems final plant populations were comparably reduced over the sole cropping. This might be due to competition for sunlight, space and nutrients between the groundnut and sweet corn in additive intercropping systems as compared to sole groundnut. In replacement intercropping systems final plant populations were comparably reduced over the sole groundnut due to groundnut row replaced by intercrop based on row proportions
(Konlan et al., 2013 and
Baishya et al., 2021).
Significantly lowest number of branches per plant, reduced leaf area, dry matter as well as leaf area index of groundnut was recorded under intercropping systems of groundnut + sweet corn (1:1) in additive series (T
6) might be severe competition for resources by intercrop with groundnut, which made poor utilization of sunlight, space and nutrients between the groundnut and sweet corn in the intercropping system, consequently growth was suppressed owing to reduced photosynthesis and partitioning of photosynthates (
Polthanee and Vidhaya, 2003;
Ghosh, 2004;
Inal et al., 2007; Mandal et al., 2014 and
Konlan et al., 2015). Sweet corn has a significant effect on groundnut nitrogen fixation traits in intercropping systems
(Zhang et al., 2024). Poor nodulation in groundnuts may be the result of less light owing to shade
(Nambiar et al., 1982). By limiting photosynthesis and the energy supply to roots, this decreased light energy impacts N2 fixation. This lowers nodulation and nodule size, which in turn results in lower fresh weight of root nodules per plant in intercropping regimens.
The height of the plant was significantly highest under intercropping systems of groundnut + sweet corn (1:1) in additive series (T6) might be due to shading imposed on the groundnut by the sweet corn leads to lengthening of the groundnut internodes in order to capture sunlight. The taller groundnut plants observed in the groundnut-sweet corn association were probably a consequence of light and space competition with the sweet corn (
Musthapha et al., 2016).
Effect of intercropping on yield attributes and yield
An appraisal of data on yield attributes and yield are presented in Table 2 indicated that which was significantly influenced by various mix/intercropping systems in pooled results of both the year of 2020-21 and 2021-22.
The sole groundnut (T
1) had significantly the highest 100-kernel weight (41.0 g), shelling percentage (70.32%), number of pods/plant (20.9), pod yield (1831 kg/ha) and haulm yield (3478 kg/ha) and which was remain statistically at par with treatments intercropping of groundnut + sweet corn (2:1) in replacement series (T
4), groundnut + sweet corn (3:1) in replacement series (T
5), groundnut + sweet corn (3:1) in additive series (T
8) and paired row sowing of groundnut + sweet corn (2:1) at 30-60-30 cm (T
10) (Except, treatment T4 in 100-kernel weight, treatments T4 and T8 in number of pods/plant and treatments T
4 and T
5 in pod and haulm yield) which may be due to sole crop enjoyed higher availability of nutrient, moisture, light, space
etc. and improved yield attributes and finally gave higher crop yield
(Baishya et al., 2021). Overall yield reduction of groundnut was observed under mix/intercropping situation when compared with sole cropping. Whereas, harvest index (35.21), groundnut pod equivalent yield (3221 kg/ha), land equivalent ratio (1.88) and area time equivalent ratio (1.75) were recorded significantly the highest under paired row sowing of groundnut + sweet corn (2:1) at 30-60-30 cm (T
10) and which was remain statistically at par with treatments T8 in pooled results (Table 2). The land equivalent ratio of 1.88 signifies that a sole cropping system would need 88% more space to achieve the yield of an intercropping system. Additionally, it indicates that mix/intercropping systems are more biologically sustainable than sole cropping. Because area time equivalent ratio takes into account the variation in time utilized by the component crops of different mix/intercropping systems, it offers a more realistic comparison of the yield benefit of intercropping over that of sole cropping than does LER. The groundnut + sweet corn intercropping system’s ATER was greater than one under all treatments (treatments T
2, T
3 and T
6) indicating the benefit of mix/intercropping systems. On the other hand, the ATER was less than one, indicating that the intercropping system did not maximize the use of all resources
(Kathmale et al., 2014; Rajaii and Dahmardeh, 2014;
Sutharsan and Srikrishnah, 2015;
Musthapha et al., 2016; Kithan and Longkumer, 2017;
Telkar et al., 2018; Chovatia et al., 2020 and
Baishya et al., 2021).
The presence of groundnut in the paired row system probably had more synergistic effect as compared to antagonistic effect as well as more space available to groundnut plant which increase photosynthetic activities and resulted in higher growth and yield attributes which ultimately reflected in yield of groundnut (
Jana and Saren, 1998;
Polthanee and Vidhaya, 2003;
Ghosh, 200,
Bhagat et al., 2007; Dutta and Bandyopadhyay, 2006;
Konlan et al., 2013; Mandal et al., 2014;
Shwethanjali et al., 2018; Sepat et al., 2019; Das et al., 2020; Baishya et al., 2021 and
Abba et al., 2024).
The relationship between the pod yield of groundnut and other important growth, yield attributes and haulm yield has been studied. The data on correlation (r) are graphically shown in Fig 1 and 2.
Result revealed that growth attributes
viz., number of root nodules per plant and yield attributes
viz., 100-kernel weight and number of pods per plant and haulm yield of groundnut showed positive and highly significant correlation with pod yield of groundnut. While plant height at harvest shown their negative correlation with pod yield in pooled result.