Ecological Weed Management Practices in Maize (Zea mays L.): A Review

R
R. Chethan1
B
B. Sandhya Rani2,*
G
G. Karuna Sagar3
1Department of Agronomy, S.V. Agricultural College, Acharya N.G. Ranga Agricultural University, Tirupati-517 502, Andhra Pradesh, India.
2Department of Agronomy, Regional Agricultural Research Station, Tirupati-517 502, Andhra Pradesh, India.
3University Librarian, Acharya N.G. Ranga Agricultural University, Lam, Guntur-522 034, Andhra Pradesh, India.

In India the highest yield loss (34%) is caused by weeds, followed by insects (29%), pathogens (22%), storage pests (7%) and others (8%). Weeds compete with the crop for both above and below ground resources. Weed infestation is a serious challenge for maize growers in our country. Maize is sensitive to weeds especially during early stages of development and thus weed infestation from germination to 45 DAS causes maximum reduction in yield. Herbicides are efficient tools for checking weed infestation and its usage is increasing throughout the globe due to increasing labour cost and quick weed control in various situations. Continuous non-judicious use of herbicides for weed management leads to loss of biodiversity, environmental pollution and also develops herbicide resistance in weeds. Weed persistence is more in organic farming due to the extensive usage of organic manures, which act as weed seed reservoirs.

Maize (Zea mays L.) is the most versatile crop with wider adaptability to varied agro-ecological regions and diverse growing seasons. Besides serving as human food and animal feed, the importance of this crop also lies in its wide industrial applications. Maize is grown on 194 million ha area, in more than 170 countries across the globe with 1148 million metric tonns of production and with a productivity of 5755 kg ha-1. In India during 2021 it is grown on 9.89 million ha area with 31.65 million tonnes of production and with a productivity of 3199 kg ha-1 (www.indiastat.com).
       
Among various biotic (insect, pest, predators, weed, etc.) and a biotic factors (drought, salinity, heat, etc.) that hinder maize production, weed is considered among the foremost factors restricting the maize crop yield (Neelam and Maneesha, 2022). Corn is sensitive to weeds especially during early stages of development and thus weed infestation from germination to 45 days after sowing (DAS) causes maximum reduction in yield and the extent of yield losses vary from 28-100% depending upon the intensity, nature and duration. Weeds not only decrease the crop yield but also harbour insect-pest and diseases and in some cases, they serve as an alternate host for pests.
       
Herbicide usage is a key component in almost all weed management strategies, but indiscriminate use of these herbicides has resulted in serious ecological and environ-mental problems. Presently, there are 58 weed species in corn which are resistant to herbicides. Weed persistence is more in organic farming due to the extensive usage of organic manures- which act as weed seed reservoirs, mulches, biomass which exacerbates the weed multiplication and growth (Sanbagavalli et al., 2020). Keeping in view the importance of weed management in organic crop production a brief review is presented on the nature of weed spectrum in maize, competition between crops and weeds and different organic weed management practices.

Weed flora associated with maize
 
The predominant weed species observed in the experimental field of maize were Cyperus rotundus, Digitaria Sanguinalis, Dactyloctenium aegyptium, Blainvillea acmella, Lagascea mollis and Commelina benghalensis on sandy loam soils of Tirupati andhra Pradesh (Saimaheswari et al., 2022). Krishnaprabu (2020) reported that predominant weed species observed in the maize experimental field were Ageratum conyzoides, Alternanthera Phyloxeroides, Bidens pilosa, Borreria hispida, Galinsoga parviflora and Spilanthesacmella among the broadleaved weeds, Cynodondactylon, Digitaria marginata, Digitaria sanguinalis, Panicum repens and Eleusine indica among grasses and Cyperus rotundus and Fimbristylis miliacea among sedges in sandy clay loam soils of Annamalainagar, Tamil Nadu. Lavanya et al., (2020) observed that the most dominant weeds in maize were Cynodon dactylon, Dactyloctenium aegyptium, Echinochloa colona, Amaranthus viridis, Digera arvensis and Trianthema portulacastrum in sandy loam soils of Coimbatore, Tamil Nadu. The predominant weed species associated with corn were Trianthema portulacastrum, Digera arvensis, Echinochloa colona, Digitaria longiflora, Dactyloctenium aegyptium and Parthenium hysterophorus in sandy clay loam soils of Coimbatore, Tamil Nadu (Sathyapriya and Chinnusamy, 2020). Nazir et al., (2019) observed that the major weeds associated with baby corn in Manasbal, Kashmir were Cynodon dactylon, Sorghum halepense, Poa annua, Portulaca oleracea, Convolvulus arvensis, Amaranthus viridis, Chenopodium album and Cyperus rotundus. The predominant weed species observed in the experimental field of maize were Cyperus rotundus a sedge, Cynodon dactylon among grasses and Digera arvensis, Trianthemaportulacastrum, Cleome viscosa and Phyllanthus niruri among broadleaved weeds in sandy clay loam soils of Killikulam, Tamil Nadu (Naik and Velayutham, 2018). Rajeshkumar et al., (2018) noticed grasses like Echinochloa colona, Eleusine indica and Dactyloctenium aegyptium, Cyperus rotundus a sedge and Trianthema portulacastrum, Boerhavia diffusa and Digera arvensis among the broadleaved weeds in maize in clay loam soils of Madurai, Tamil Nadu. Kumar et al., (2017) observed the infestation of Cynodon dactylon, Cyperus rotundus, Amaranthus viridis, Anagalis arvensis, Argemone mexicana, Chenopodium album, Melilotusindica, Oxaliscorniculata, Convolvulus arvensis, Rumex retroflex and Parthenium hysterophorus in maize field on sandy loam soils of Sabour, Bihar. Ram et al., (2017) reported that predominant weed species associated with maize in sandy loamy soils of Hyderabad, Telangana were Parthenium hysterophorus, Melilotus alba, Cyperus rotundus, Trianthema portulacastrum, Dactyloctenium aegyptium, Cynodon dactylon, Digera muricata, Amaranthus viridis, Commelina benghalensis, Eragrostis unioloides, Chenopodium album, Trichodesma indicum, Digitaria sanguinalis, Euphorbia geniculataand Echinochloa colona.
 
Weed competition
 
The critical period of crop weed competition varies from 2 to 7 WAS in maize (Shrestha et al., 2019). Imoloame and Omolalye (2017) observed that weed interference in maize from 3 to 6 WAS significantly depressed the growth parameters and kernel yield of maize. Das et al., (2016) reported that initial 6 WAS was found to be very susceptible to weed infestation in maize. Singh et al., (2016) noticed that in spring maize, the critical crop weed competition period starts from 30 DAS and continued up to 60 DAS. Critical period for weed removal ranged from 21 to 29 DAS in maize to prevent yield losses (Amare et al., 2014). Fahad et al., (2014) reported that infestation of weeds at every stage of crop led to significant yield loss in maize and weed competition during the early crop growth stages of maize should be low to achieve higher yields. The period of high emergence of weeds in maize was between 3 and 8 to 10 weeks after planting (Takim et al., 2014).

Yield loss due to weeds in maize
 
Shrestha et al., (2019) reported that weed infestation might led to 20 to 80 per cent reduction in maize yield. Imoloame and Omolalye (2017) observed 89 per cent yield loss in maize as a result of uncontrolled weed infestation. Kernel losses in maize varied between 28-100 per cent, if weeds were not controlled during the critical period of crop weed competition (Kumar et al., 2017). During Kharif season maize suffers from heavy weed infestation, which may cause yield losses ranging from 28-100 per cent depending upon the intensity and nature of infestation of weeds (Das et al., 2016). Weeds in maize cause substantial yield loss ranging from 18 to 85 per cent (Jagadish et al., 2016). Presence of weeds in winter maize reduced the maize yields by 27-60 per cent, depending upon the growth and persistence of weed population (Anil et al., 2015). Yakadri et al., (2015) stated that there was a yield reduction of 77, 44 and 38 per cent due to grasses, broadleaved weeds and sedges respectively in maize.
 
Effect of organic weed management practices on weed dynamics, growth, yield and economics of maize
 
Weed density and dry weight
 
Saimaheswari et al., (2022) reported that hand weeding twice at 15 and 30 DAS recorded significantly lower density and biomass of grasses, sedges, broadleaved and total weeds over the rest of treatments in maize. Masud et al., (2021) observed significantly lower weed biomass with cowpea live mulch when compared to rest of the treatments in maize. Asif et al., (2020) reported that application of saw dust mulch at 4 t ha-1 recorded significantly lower weed density and dry weight when compared to control in maize. Kaur et al., (2019) stated that live mulch with cowpea has significantly reduced the weed density and weed dry matter, which was however comparable with moong bean live mulch in maize. Two hand weedings at 20 and 40 DAS recorded significantly lowest weed density and dry weight at 30, 45 and 60 DAS in maize (Soren et al., 2018). Stanzen  et al. (2017) observed minimum density and biomass of weeds with two hand weedings at 15 and 30 DAS in maize. Foliar application of combined aqueous extracts of sorghum and sunflower each at 15 L ha-1 at 20 DAS of corn reduced the density of Trianthema portulacastrum by 46 per cent and Cyperus rotundus by 56 per cent respectively (Naeem et al., 2016). Seremesic et al. (2016) reported that pre emergence application of maize gluten at 3 t ha-1 resulted in significant reduction in weed density in maize. Arif et al., (2015) stated that combined exogenous application of sorghum, sunflower and brassica extract at 18 L ha-1 at 25 and 40 DAS significantly suppressed the weeds number and biomass in maize. Application of aqueous extract of sunflower along with sorghum and brassica aqueous extracts each at 15 L ha-1 twice at 15 and 30 DAS resulted in reduction of weed density by 74 per cent in maize (Ihsan et al., 2015). Jamshidi et al., (2013) concluded that planting of maize at a density of 9 plants m-2 and simultaneously intercropping with cowpea at a density of 30 plants m-2 effectively suppressed the weed growth and biomass by 39.6 per cent. Combined application of mixture of sorghum, sunflower and rice allelopathic water extracts each at 15 L ha-1 reduced the density of barnyard grass, flat sedge and crowfoot grass by 75, 67 and 74 per cent and their dry weight by 66, 71 and 76 per cent respectively in rice (Rehman et al., 2012). Talebbeigi and Ghadiri (2012) reported that growing cowpea as a live mulch at 22 plants m-2 with a suppression period of 75 days recorded lowest weed biomass and dry weight of weeds at harvest of maize. Corn gluten meal as pre emergence application at 300 g m-2 inhibited root elongation of many broadleaf weeds in greenhouse studies (Hebert and lyons, 2011). Pre emergence application of corn gluten meal at 4 t ha-1 provided complete control of Dactyloctenium aegyptium, Lolium multiflorum and Eleusine indica and substantial inhibition of Setaria viridis and moderate control of Sorghum halepense and Cyperus esculentus in green house studies (Abouziena  et al., 2009).
 
Weed control efficiency
 
Baldaniya et al., (2018) reported that hand weeding twice at 20 and 40 DAS recorded highest weed control efficiency compared to pre and post emergence herbicide application treatments in corn. Hand weeding at 20 and 40 DAS recorded the highest weedcontrol efficiency at all the growth stages of maize compared to rest of the weed management practices tested (Gupta et al., 2018). Enhanced weed control efficiency at all the growth stages of maize was noticed with hand weeding performed at 20 and 40 DAS (Satyendra et al., 2018). Kumar et al., (2017) observed that cowpea live mulch in maize registered highest weed control efficiency at 15, 35 and 60 DAS when compared to rest of the treatments. At physiological maturity stage of maize, significantly higher WCE was noticed in live mulch with cowpea and was at par with black and white polythene mulch (Ram et al., 2017). Webber et al., (2008) noticed that application of corn gluten meal at 4000 kg ha-1 recorded 72 per cent of total weed control and 83 per cent of broadleaf weed control up to 46 days after planting in spring transplanted corn.
 
Weed index
 
Dhivya et al., (2021) noticed that groundnut shell mulch at 5 t ha-1 + one hand weeding at 20 DAS recorded the lowest weed index when compared to intercropping with dhaincha and in-situ incorporation on 30 DAS in corn. Vidyashree et al., (2019) reported that application of mango leaves mulch between the rows of sunflower at 4 t ha-1 recorded significantly lower weed index when compared to live mulch with sunhemp at 40 kg ha-1. Hand weeding and intercultivation at 15 and 30 DAS recorded significantly minimum weed index when compared to rest of the treatments tried in popcorn (Barad et al., 2016).
 
Phytotoxicity of maize
 
Germination percentage of maize was inhibited by dry aqueous extracts of maize leaf by 7.81%, germination index by 16.51%, increased mean germination time by 25.53%, decreased plumule and radical lengths by 29.00 and 36.12% respectively and lowered maize seedling dry biomass by 34.02% (Hussain et al., 2018). Rawat et al., (2013) reported that application of sunflower extract at 15 per cent concentration twice at 15 and 30 DAS inhibited the germination and growth of Vigna radiata and Pennisetum glaucum in a laboratory experiment. The allelochemicals secreted by sunflower inhibited germination, reduced the shoot length, root length, fresh weight, dry weight and chlorophyll content and increased the values of ABA, protein, proline, sugar, superoxide dismutase and peroxidase contents in maize seedlings (Kamal, 2011). Khan et al., (2007) revealed that eucalyptus aqueous extract application at 10 g L-1 of water reduced the seed germination of maize by 12 to 18 per cent over water applied treatment in pot culture experiment. Eucalyptus boiled extract decreased seed germination by 66 per cent when compared to 99 per cent germination in the control treatment in maize.
 
Growth parameters of maize
 
Masud et al., (2021) observed significantly tallest plants in maize by live mulch with cowpea when compared to rest of the treatments. Application of saw dust mulch at 4 t ha-1 recorded significantly higher plant height when compared to control in maize (Asif et al., 2020). Bako et al., (2020) recorded significantly highest plant height, stem girth, number of leaves plant-1 and leaf area index with application of groundnut shells at 7.5 t ha-1 when compared to rest of the treatments in maize. Hand weeding twice at 20 and 40 DAS recorded significantly higher plant height, leaf area index and dry matter production in maize (Kandasamy, 2018). Saw dust mulch at 4 t ha-1 recorded higher number of leaves plant-1, plant height, stem girth, leaf area index and dry matter production in maize (Wuese  et al., 2018). Hand weeding twice at 20 and 40 DAS resulted in maximum plant height, leaf area index and dry matter production in maize (Ram et al., 2017). Foliar application of aqueous plant extracts of sorghum and sunflower each at 15 L ha-1 at 20 DAS increased the dry matter production of maize by 69 per cent when compared to control (Naeem et al., 2016). Mathukia  et al. (2015) noticed significantly highest plant height, leaf area index, dry matter production, plant spread and number of branches plant-1 with the groundnut shells mulch at 5 t ha-1, which was comparable with wheat straw mulch at 5 t ha-1. Ihsan et al., (2015) noticed that application of sunflower plant water extract along with sorghum and brassica aqueous extracts each at 18 L ha-1 twice at 15 and 30 DAS resulted in increased plant height by 20 per cent in maize.
 
Yield attributes and yield of maize
 
Masud et al., (2021) observed increased kernel and stover yield with cowpea live mulch when compared to rest of the treatments in maize. Asif et al., (2020) reported that application of saw dust mulch at 4 t ha-1 recorded significantly higher number of kernels cob-1, kernel weight cob-1, 100 kernel weight, kernel yield, stover yield and harvest index in maize. Bako et al., (2020) recorded significantly higher cob length, cob girth, number of kernels cob-1, 100 kernel weight, stover yield, kernel yield and harvest index of maize with application of groundnut shells as mulch at 7.5 t ha-1 when compared to rest of the treatments. Khan et al., (2020) reported significantly higher kernel weight cob-1, number of kernels cob-1, kernel yield and harvest index of maize with hand weeding twice at 20 and 40 DAS in maize. Mahto et al., (2020) revealed that significantly higher cob length, cob girth, number of kernel rows cob-1, 100 kernel weight and kernel yield of maize was recorded with hand weeding twice at 20 and 40 DAS. Hand weeding twice at 15 and 30 DAS recorded higher cob length, cob girth, kernel rows cob-1, number of kernels cob-1, 100 kernel weight, kernel yield and stover yield when compared to other treatments in maize (Rani et al., 2019). Prithwiraj et al., (2018) reported highest green cob yield of maize with hand weeding twice at 20 and 40 DAS.
       
Higher cob length, cob diameter, number of kernels row-1, number of kernels cob-1, test weight and kernel yield of maize was recorded under maize + cowpea as live mulch treatment (Kumar et al., 2017). Kurre et al., (2017) recorded highest kernel yield of maize with hand weeding twice at 20 and 40 DAS when compared to rest of the treatments. Parameswari et al., (2017) reported that the number of kernels cob-1, kernel weight cob-1, test weight, stover yield and kernel yield were found to be significantly higher with hand weeding twice at 20 and 40 DAS in maize. Ram et al., (2017) observed significantly higher kernel yield in live mulch with cowpea in maize when compared to weedy check. Triveni et al., (2017) noticed significantly higher cob length, cob girth, number of kernel rows cob-1, test weight, kernel yield and stover yield with hand weeding twice at 20 and 40 DAS in maize. Sraw  et al. (2016) reported that higher cob length, cob girth, number of kernel rows cob-1, number of kernels cob-1 and kernel yield of maize was recorded in live mulch with cowpea as compared to rest of the treatments tried. Samanth et al., (2015) reported higher cob length, cob weight, number of kernels cob-1, 100 kernel weight and kernel yield of maize with hand weeding twice at 20 and 40 DAS. Higher kernel yield of maize was recorded with hand weeding twice at 20 and 40 DAS when compared to rest of the treatments tried (Dash and Mishra, 2014). Jamshidi et al., (2013) observed that under weed infested conditions, intercropping of maize with cowpea as a live mulch increased kernel yield of maize. Talebbeigi and Ghadiri (2012) reported higher number of kernel rows cob-1, number of kernels row-1, number of kernels cob-1, test weight and kernel yield with cowpea live mulch at 22 plants m-2 in maize.
 
Nutrient uptake by maize crop  
 
Two hand weedings at 15 and 30 DAS recorded significantly higher total N, P and K uptake by sweet corn when compared to rest of the treatments (Bhalerao et al., 2020). Hiremath et al., (2020) reported that live mulch of sunhemp with maize in 1:2 ratio recorded significantly higher uptake of N, P and K by maize. Maximum uptake of N, P and K by maize was recorded with two hand weedings at 15 and 30 DAS when compared to rest of the treatments (Jaybhaye et al., 2019). Yadav et al., (2018) found that highest uptake of N, P and K by kernel and stover of maize was recorded with two hand weedings at 20 and 40 DAS. Hand weeding twice at 15 and 30 DAS recorded significantly highest uptake of nutrients by maize when compared to rest of the treatments (Lakshmi and Martin, 2017). Samant et al., (2015) concluded that maximum uptake of N, P and K by maize was observed with two hand weedings at 20 and 40 DAS. Among weed management practices, highest N, P and K removal by kernel and stover of winter maize was noticed with hand weeding twice at 15 and 30 DAS (Kour et al., 2014).
 
Nutrient uptake by weeds in maize
 
Live mulch with cowpea up to 50 DAS recorded significantly lowest N, P and K uptake by weeds at harvest of maize (Choudhary and Ezung, 2020). Jaybhaye et al., (2019) observed that two hand weedings at 15 and 30 DAS recorded significantly lowest nutrients removal by weeds in maize. Swetha et al., (2018) reported that nitrogen, phosphorus and potassium uptake by weeds at harvest was significantly lower in hand weeding at 20 and 40 DAS when compared to rest of the treatments in maize.Hand weeding twice at 15 and 30 DAS recorded significantly the lowest uptake of nutrients by weeds when compared to rest of the treatments in maize (Lakshmi and Martin, 2017). Stanzen et al., (2017) recorded significantly lowest N, P and K uptake by weeds with two hand weedings at 15 and 30 DAS in maize. Samant et al., (2015) concluded that minimum uptake of N, P and K by weeds was observed with two hand weedings at 20 and 40 DAS in maize.Among all the weed management practices, the uptake of N, P and K by weeds in maize was found to be significantly lowest in hand weeding twice at 15 and 30 DAS (Kour et al., 2014).
 
Economics
 
Hiremath et al., (2020) reported that live mulch of sunhemp with maize in 1:2 ratio recorded significantly higher net returns and B:C ratio in maize. Dutta et al., (2016) recorded maximum net returns and B:C ratio with hand weeding twice at 20 and 40 DAS in maize. Maize intercropped with cowpea as a live mulch upto 30 DAS recorded maximum net returns and B:C ratio when compared to control (Sraw et al., 2016). Post emergence combined application of aqueous extracts of sorghum+ sunflower+ brassica each at 18 l ha-1 at 25 and 40 DAS gave higher net returns in maize (Arif et al., 2015). Samant et al., (2015) recorded maximum gross returns, net returns and B:C ratio withtwo hand weedings at 20 and 40 DAS in maize. Singh et al., (2015) noticed significantly highest gross returns, net returns and B:C ratio with cowpea as a live mulch in maize. Maximum net returns and B:C ratio were realized with hand weeding and intercultivation at 15 and 30 DAS in sweet corn (Dobaria et al., 2014). The maximum gross returns was obtained with hand weeding twice at 20 and 40 DAS in maize when compared to rest of the treatments (Sanodiya et al., 2013).
Weeds have a more direct influence on yields of crops than any other pest in developing countries like India. Weeds not only cause severe crop losses but also compete with farmers and their families to spend a considerable amount and their time on weeding. Different weed control methods have been used to manage the weeds but the usage of herbicides is restricted in organic farming. In organic farming weeds can be effectively managed by mechanical weeding, Hand hoeing, Hand weeding, application of organic mulches, growing of live mulches, inter cropping and  cover crops.  Apart from the above allelopathy and mycoherbicides can play an important role in reducing the weed infestation.
All authors declare that they have no conflicts of interest.

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Ecological Weed Management Practices in Maize (Zea mays L.): A Review

R
R. Chethan1
B
B. Sandhya Rani2,*
G
G. Karuna Sagar3
1Department of Agronomy, S.V. Agricultural College, Acharya N.G. Ranga Agricultural University, Tirupati-517 502, Andhra Pradesh, India.
2Department of Agronomy, Regional Agricultural Research Station, Tirupati-517 502, Andhra Pradesh, India.
3University Librarian, Acharya N.G. Ranga Agricultural University, Lam, Guntur-522 034, Andhra Pradesh, India.

In India the highest yield loss (34%) is caused by weeds, followed by insects (29%), pathogens (22%), storage pests (7%) and others (8%). Weeds compete with the crop for both above and below ground resources. Weed infestation is a serious challenge for maize growers in our country. Maize is sensitive to weeds especially during early stages of development and thus weed infestation from germination to 45 DAS causes maximum reduction in yield. Herbicides are efficient tools for checking weed infestation and its usage is increasing throughout the globe due to increasing labour cost and quick weed control in various situations. Continuous non-judicious use of herbicides for weed management leads to loss of biodiversity, environmental pollution and also develops herbicide resistance in weeds. Weed persistence is more in organic farming due to the extensive usage of organic manures, which act as weed seed reservoirs.

Maize (Zea mays L.) is the most versatile crop with wider adaptability to varied agro-ecological regions and diverse growing seasons. Besides serving as human food and animal feed, the importance of this crop also lies in its wide industrial applications. Maize is grown on 194 million ha area, in more than 170 countries across the globe with 1148 million metric tonns of production and with a productivity of 5755 kg ha-1. In India during 2021 it is grown on 9.89 million ha area with 31.65 million tonnes of production and with a productivity of 3199 kg ha-1 (www.indiastat.com).
       
Among various biotic (insect, pest, predators, weed, etc.) and a biotic factors (drought, salinity, heat, etc.) that hinder maize production, weed is considered among the foremost factors restricting the maize crop yield (Neelam and Maneesha, 2022). Corn is sensitive to weeds especially during early stages of development and thus weed infestation from germination to 45 days after sowing (DAS) causes maximum reduction in yield and the extent of yield losses vary from 28-100% depending upon the intensity, nature and duration. Weeds not only decrease the crop yield but also harbour insect-pest and diseases and in some cases, they serve as an alternate host for pests.
       
Herbicide usage is a key component in almost all weed management strategies, but indiscriminate use of these herbicides has resulted in serious ecological and environ-mental problems. Presently, there are 58 weed species in corn which are resistant to herbicides. Weed persistence is more in organic farming due to the extensive usage of organic manures- which act as weed seed reservoirs, mulches, biomass which exacerbates the weed multiplication and growth (Sanbagavalli et al., 2020). Keeping in view the importance of weed management in organic crop production a brief review is presented on the nature of weed spectrum in maize, competition between crops and weeds and different organic weed management practices.

Weed flora associated with maize
 
The predominant weed species observed in the experimental field of maize were Cyperus rotundus, Digitaria Sanguinalis, Dactyloctenium aegyptium, Blainvillea acmella, Lagascea mollis and Commelina benghalensis on sandy loam soils of Tirupati andhra Pradesh (Saimaheswari et al., 2022). Krishnaprabu (2020) reported that predominant weed species observed in the maize experimental field were Ageratum conyzoides, Alternanthera Phyloxeroides, Bidens pilosa, Borreria hispida, Galinsoga parviflora and Spilanthesacmella among the broadleaved weeds, Cynodondactylon, Digitaria marginata, Digitaria sanguinalis, Panicum repens and Eleusine indica among grasses and Cyperus rotundus and Fimbristylis miliacea among sedges in sandy clay loam soils of Annamalainagar, Tamil Nadu. Lavanya et al., (2020) observed that the most dominant weeds in maize were Cynodon dactylon, Dactyloctenium aegyptium, Echinochloa colona, Amaranthus viridis, Digera arvensis and Trianthema portulacastrum in sandy loam soils of Coimbatore, Tamil Nadu. The predominant weed species associated with corn were Trianthema portulacastrum, Digera arvensis, Echinochloa colona, Digitaria longiflora, Dactyloctenium aegyptium and Parthenium hysterophorus in sandy clay loam soils of Coimbatore, Tamil Nadu (Sathyapriya and Chinnusamy, 2020). Nazir et al., (2019) observed that the major weeds associated with baby corn in Manasbal, Kashmir were Cynodon dactylon, Sorghum halepense, Poa annua, Portulaca oleracea, Convolvulus arvensis, Amaranthus viridis, Chenopodium album and Cyperus rotundus. The predominant weed species observed in the experimental field of maize were Cyperus rotundus a sedge, Cynodon dactylon among grasses and Digera arvensis, Trianthemaportulacastrum, Cleome viscosa and Phyllanthus niruri among broadleaved weeds in sandy clay loam soils of Killikulam, Tamil Nadu (Naik and Velayutham, 2018). Rajeshkumar et al., (2018) noticed grasses like Echinochloa colona, Eleusine indica and Dactyloctenium aegyptium, Cyperus rotundus a sedge and Trianthema portulacastrum, Boerhavia diffusa and Digera arvensis among the broadleaved weeds in maize in clay loam soils of Madurai, Tamil Nadu. Kumar et al., (2017) observed the infestation of Cynodon dactylon, Cyperus rotundus, Amaranthus viridis, Anagalis arvensis, Argemone mexicana, Chenopodium album, Melilotusindica, Oxaliscorniculata, Convolvulus arvensis, Rumex retroflex and Parthenium hysterophorus in maize field on sandy loam soils of Sabour, Bihar. Ram et al., (2017) reported that predominant weed species associated with maize in sandy loamy soils of Hyderabad, Telangana were Parthenium hysterophorus, Melilotus alba, Cyperus rotundus, Trianthema portulacastrum, Dactyloctenium aegyptium, Cynodon dactylon, Digera muricata, Amaranthus viridis, Commelina benghalensis, Eragrostis unioloides, Chenopodium album, Trichodesma indicum, Digitaria sanguinalis, Euphorbia geniculataand Echinochloa colona.
 
Weed competition
 
The critical period of crop weed competition varies from 2 to 7 WAS in maize (Shrestha et al., 2019). Imoloame and Omolalye (2017) observed that weed interference in maize from 3 to 6 WAS significantly depressed the growth parameters and kernel yield of maize. Das et al., (2016) reported that initial 6 WAS was found to be very susceptible to weed infestation in maize. Singh et al., (2016) noticed that in spring maize, the critical crop weed competition period starts from 30 DAS and continued up to 60 DAS. Critical period for weed removal ranged from 21 to 29 DAS in maize to prevent yield losses (Amare et al., 2014). Fahad et al., (2014) reported that infestation of weeds at every stage of crop led to significant yield loss in maize and weed competition during the early crop growth stages of maize should be low to achieve higher yields. The period of high emergence of weeds in maize was between 3 and 8 to 10 weeks after planting (Takim et al., 2014).

Yield loss due to weeds in maize
 
Shrestha et al., (2019) reported that weed infestation might led to 20 to 80 per cent reduction in maize yield. Imoloame and Omolalye (2017) observed 89 per cent yield loss in maize as a result of uncontrolled weed infestation. Kernel losses in maize varied between 28-100 per cent, if weeds were not controlled during the critical period of crop weed competition (Kumar et al., 2017). During Kharif season maize suffers from heavy weed infestation, which may cause yield losses ranging from 28-100 per cent depending upon the intensity and nature of infestation of weeds (Das et al., 2016). Weeds in maize cause substantial yield loss ranging from 18 to 85 per cent (Jagadish et al., 2016). Presence of weeds in winter maize reduced the maize yields by 27-60 per cent, depending upon the growth and persistence of weed population (Anil et al., 2015). Yakadri et al., (2015) stated that there was a yield reduction of 77, 44 and 38 per cent due to grasses, broadleaved weeds and sedges respectively in maize.
 
Effect of organic weed management practices on weed dynamics, growth, yield and economics of maize
 
Weed density and dry weight
 
Saimaheswari et al., (2022) reported that hand weeding twice at 15 and 30 DAS recorded significantly lower density and biomass of grasses, sedges, broadleaved and total weeds over the rest of treatments in maize. Masud et al., (2021) observed significantly lower weed biomass with cowpea live mulch when compared to rest of the treatments in maize. Asif et al., (2020) reported that application of saw dust mulch at 4 t ha-1 recorded significantly lower weed density and dry weight when compared to control in maize. Kaur et al., (2019) stated that live mulch with cowpea has significantly reduced the weed density and weed dry matter, which was however comparable with moong bean live mulch in maize. Two hand weedings at 20 and 40 DAS recorded significantly lowest weed density and dry weight at 30, 45 and 60 DAS in maize (Soren et al., 2018). Stanzen  et al. (2017) observed minimum density and biomass of weeds with two hand weedings at 15 and 30 DAS in maize. Foliar application of combined aqueous extracts of sorghum and sunflower each at 15 L ha-1 at 20 DAS of corn reduced the density of Trianthema portulacastrum by 46 per cent and Cyperus rotundus by 56 per cent respectively (Naeem et al., 2016). Seremesic et al. (2016) reported that pre emergence application of maize gluten at 3 t ha-1 resulted in significant reduction in weed density in maize. Arif et al., (2015) stated that combined exogenous application of sorghum, sunflower and brassica extract at 18 L ha-1 at 25 and 40 DAS significantly suppressed the weeds number and biomass in maize. Application of aqueous extract of sunflower along with sorghum and brassica aqueous extracts each at 15 L ha-1 twice at 15 and 30 DAS resulted in reduction of weed density by 74 per cent in maize (Ihsan et al., 2015). Jamshidi et al., (2013) concluded that planting of maize at a density of 9 plants m-2 and simultaneously intercropping with cowpea at a density of 30 plants m-2 effectively suppressed the weed growth and biomass by 39.6 per cent. Combined application of mixture of sorghum, sunflower and rice allelopathic water extracts each at 15 L ha-1 reduced the density of barnyard grass, flat sedge and crowfoot grass by 75, 67 and 74 per cent and their dry weight by 66, 71 and 76 per cent respectively in rice (Rehman et al., 2012). Talebbeigi and Ghadiri (2012) reported that growing cowpea as a live mulch at 22 plants m-2 with a suppression period of 75 days recorded lowest weed biomass and dry weight of weeds at harvest of maize. Corn gluten meal as pre emergence application at 300 g m-2 inhibited root elongation of many broadleaf weeds in greenhouse studies (Hebert and lyons, 2011). Pre emergence application of corn gluten meal at 4 t ha-1 provided complete control of Dactyloctenium aegyptium, Lolium multiflorum and Eleusine indica and substantial inhibition of Setaria viridis and moderate control of Sorghum halepense and Cyperus esculentus in green house studies (Abouziena  et al., 2009).
 
Weed control efficiency
 
Baldaniya et al., (2018) reported that hand weeding twice at 20 and 40 DAS recorded highest weed control efficiency compared to pre and post emergence herbicide application treatments in corn. Hand weeding at 20 and 40 DAS recorded the highest weedcontrol efficiency at all the growth stages of maize compared to rest of the weed management practices tested (Gupta et al., 2018). Enhanced weed control efficiency at all the growth stages of maize was noticed with hand weeding performed at 20 and 40 DAS (Satyendra et al., 2018). Kumar et al., (2017) observed that cowpea live mulch in maize registered highest weed control efficiency at 15, 35 and 60 DAS when compared to rest of the treatments. At physiological maturity stage of maize, significantly higher WCE was noticed in live mulch with cowpea and was at par with black and white polythene mulch (Ram et al., 2017). Webber et al., (2008) noticed that application of corn gluten meal at 4000 kg ha-1 recorded 72 per cent of total weed control and 83 per cent of broadleaf weed control up to 46 days after planting in spring transplanted corn.
 
Weed index
 
Dhivya et al., (2021) noticed that groundnut shell mulch at 5 t ha-1 + one hand weeding at 20 DAS recorded the lowest weed index when compared to intercropping with dhaincha and in-situ incorporation on 30 DAS in corn. Vidyashree et al., (2019) reported that application of mango leaves mulch between the rows of sunflower at 4 t ha-1 recorded significantly lower weed index when compared to live mulch with sunhemp at 40 kg ha-1. Hand weeding and intercultivation at 15 and 30 DAS recorded significantly minimum weed index when compared to rest of the treatments tried in popcorn (Barad et al., 2016).
 
Phytotoxicity of maize
 
Germination percentage of maize was inhibited by dry aqueous extracts of maize leaf by 7.81%, germination index by 16.51%, increased mean germination time by 25.53%, decreased plumule and radical lengths by 29.00 and 36.12% respectively and lowered maize seedling dry biomass by 34.02% (Hussain et al., 2018). Rawat et al., (2013) reported that application of sunflower extract at 15 per cent concentration twice at 15 and 30 DAS inhibited the germination and growth of Vigna radiata and Pennisetum glaucum in a laboratory experiment. The allelochemicals secreted by sunflower inhibited germination, reduced the shoot length, root length, fresh weight, dry weight and chlorophyll content and increased the values of ABA, protein, proline, sugar, superoxide dismutase and peroxidase contents in maize seedlings (Kamal, 2011). Khan et al., (2007) revealed that eucalyptus aqueous extract application at 10 g L-1 of water reduced the seed germination of maize by 12 to 18 per cent over water applied treatment in pot culture experiment. Eucalyptus boiled extract decreased seed germination by 66 per cent when compared to 99 per cent germination in the control treatment in maize.
 
Growth parameters of maize
 
Masud et al., (2021) observed significantly tallest plants in maize by live mulch with cowpea when compared to rest of the treatments. Application of saw dust mulch at 4 t ha-1 recorded significantly higher plant height when compared to control in maize (Asif et al., 2020). Bako et al., (2020) recorded significantly highest plant height, stem girth, number of leaves plant-1 and leaf area index with application of groundnut shells at 7.5 t ha-1 when compared to rest of the treatments in maize. Hand weeding twice at 20 and 40 DAS recorded significantly higher plant height, leaf area index and dry matter production in maize (Kandasamy, 2018). Saw dust mulch at 4 t ha-1 recorded higher number of leaves plant-1, plant height, stem girth, leaf area index and dry matter production in maize (Wuese  et al., 2018). Hand weeding twice at 20 and 40 DAS resulted in maximum plant height, leaf area index and dry matter production in maize (Ram et al., 2017). Foliar application of aqueous plant extracts of sorghum and sunflower each at 15 L ha-1 at 20 DAS increased the dry matter production of maize by 69 per cent when compared to control (Naeem et al., 2016). Mathukia  et al. (2015) noticed significantly highest plant height, leaf area index, dry matter production, plant spread and number of branches plant-1 with the groundnut shells mulch at 5 t ha-1, which was comparable with wheat straw mulch at 5 t ha-1. Ihsan et al., (2015) noticed that application of sunflower plant water extract along with sorghum and brassica aqueous extracts each at 18 L ha-1 twice at 15 and 30 DAS resulted in increased plant height by 20 per cent in maize.
 
Yield attributes and yield of maize
 
Masud et al., (2021) observed increased kernel and stover yield with cowpea live mulch when compared to rest of the treatments in maize. Asif et al., (2020) reported that application of saw dust mulch at 4 t ha-1 recorded significantly higher number of kernels cob-1, kernel weight cob-1, 100 kernel weight, kernel yield, stover yield and harvest index in maize. Bako et al., (2020) recorded significantly higher cob length, cob girth, number of kernels cob-1, 100 kernel weight, stover yield, kernel yield and harvest index of maize with application of groundnut shells as mulch at 7.5 t ha-1 when compared to rest of the treatments. Khan et al., (2020) reported significantly higher kernel weight cob-1, number of kernels cob-1, kernel yield and harvest index of maize with hand weeding twice at 20 and 40 DAS in maize. Mahto et al., (2020) revealed that significantly higher cob length, cob girth, number of kernel rows cob-1, 100 kernel weight and kernel yield of maize was recorded with hand weeding twice at 20 and 40 DAS. Hand weeding twice at 15 and 30 DAS recorded higher cob length, cob girth, kernel rows cob-1, number of kernels cob-1, 100 kernel weight, kernel yield and stover yield when compared to other treatments in maize (Rani et al., 2019). Prithwiraj et al., (2018) reported highest green cob yield of maize with hand weeding twice at 20 and 40 DAS.
       
Higher cob length, cob diameter, number of kernels row-1, number of kernels cob-1, test weight and kernel yield of maize was recorded under maize + cowpea as live mulch treatment (Kumar et al., 2017). Kurre et al., (2017) recorded highest kernel yield of maize with hand weeding twice at 20 and 40 DAS when compared to rest of the treatments. Parameswari et al., (2017) reported that the number of kernels cob-1, kernel weight cob-1, test weight, stover yield and kernel yield were found to be significantly higher with hand weeding twice at 20 and 40 DAS in maize. Ram et al., (2017) observed significantly higher kernel yield in live mulch with cowpea in maize when compared to weedy check. Triveni et al., (2017) noticed significantly higher cob length, cob girth, number of kernel rows cob-1, test weight, kernel yield and stover yield with hand weeding twice at 20 and 40 DAS in maize. Sraw  et al. (2016) reported that higher cob length, cob girth, number of kernel rows cob-1, number of kernels cob-1 and kernel yield of maize was recorded in live mulch with cowpea as compared to rest of the treatments tried. Samanth et al., (2015) reported higher cob length, cob weight, number of kernels cob-1, 100 kernel weight and kernel yield of maize with hand weeding twice at 20 and 40 DAS. Higher kernel yield of maize was recorded with hand weeding twice at 20 and 40 DAS when compared to rest of the treatments tried (Dash and Mishra, 2014). Jamshidi et al., (2013) observed that under weed infested conditions, intercropping of maize with cowpea as a live mulch increased kernel yield of maize. Talebbeigi and Ghadiri (2012) reported higher number of kernel rows cob-1, number of kernels row-1, number of kernels cob-1, test weight and kernel yield with cowpea live mulch at 22 plants m-2 in maize.
 
Nutrient uptake by maize crop  
 
Two hand weedings at 15 and 30 DAS recorded significantly higher total N, P and K uptake by sweet corn when compared to rest of the treatments (Bhalerao et al., 2020). Hiremath et al., (2020) reported that live mulch of sunhemp with maize in 1:2 ratio recorded significantly higher uptake of N, P and K by maize. Maximum uptake of N, P and K by maize was recorded with two hand weedings at 15 and 30 DAS when compared to rest of the treatments (Jaybhaye et al., 2019). Yadav et al., (2018) found that highest uptake of N, P and K by kernel and stover of maize was recorded with two hand weedings at 20 and 40 DAS. Hand weeding twice at 15 and 30 DAS recorded significantly highest uptake of nutrients by maize when compared to rest of the treatments (Lakshmi and Martin, 2017). Samant et al., (2015) concluded that maximum uptake of N, P and K by maize was observed with two hand weedings at 20 and 40 DAS. Among weed management practices, highest N, P and K removal by kernel and stover of winter maize was noticed with hand weeding twice at 15 and 30 DAS (Kour et al., 2014).
 
Nutrient uptake by weeds in maize
 
Live mulch with cowpea up to 50 DAS recorded significantly lowest N, P and K uptake by weeds at harvest of maize (Choudhary and Ezung, 2020). Jaybhaye et al., (2019) observed that two hand weedings at 15 and 30 DAS recorded significantly lowest nutrients removal by weeds in maize. Swetha et al., (2018) reported that nitrogen, phosphorus and potassium uptake by weeds at harvest was significantly lower in hand weeding at 20 and 40 DAS when compared to rest of the treatments in maize.Hand weeding twice at 15 and 30 DAS recorded significantly the lowest uptake of nutrients by weeds when compared to rest of the treatments in maize (Lakshmi and Martin, 2017). Stanzen et al., (2017) recorded significantly lowest N, P and K uptake by weeds with two hand weedings at 15 and 30 DAS in maize. Samant et al., (2015) concluded that minimum uptake of N, P and K by weeds was observed with two hand weedings at 20 and 40 DAS in maize.Among all the weed management practices, the uptake of N, P and K by weeds in maize was found to be significantly lowest in hand weeding twice at 15 and 30 DAS (Kour et al., 2014).
 
Economics
 
Hiremath et al., (2020) reported that live mulch of sunhemp with maize in 1:2 ratio recorded significantly higher net returns and B:C ratio in maize. Dutta et al., (2016) recorded maximum net returns and B:C ratio with hand weeding twice at 20 and 40 DAS in maize. Maize intercropped with cowpea as a live mulch upto 30 DAS recorded maximum net returns and B:C ratio when compared to control (Sraw et al., 2016). Post emergence combined application of aqueous extracts of sorghum+ sunflower+ brassica each at 18 l ha-1 at 25 and 40 DAS gave higher net returns in maize (Arif et al., 2015). Samant et al., (2015) recorded maximum gross returns, net returns and B:C ratio withtwo hand weedings at 20 and 40 DAS in maize. Singh et al., (2015) noticed significantly highest gross returns, net returns and B:C ratio with cowpea as a live mulch in maize. Maximum net returns and B:C ratio were realized with hand weeding and intercultivation at 15 and 30 DAS in sweet corn (Dobaria et al., 2014). The maximum gross returns was obtained with hand weeding twice at 20 and 40 DAS in maize when compared to rest of the treatments (Sanodiya et al., 2013).
Weeds have a more direct influence on yields of crops than any other pest in developing countries like India. Weeds not only cause severe crop losses but also compete with farmers and their families to spend a considerable amount and their time on weeding. Different weed control methods have been used to manage the weeds but the usage of herbicides is restricted in organic farming. In organic farming weeds can be effectively managed by mechanical weeding, Hand hoeing, Hand weeding, application of organic mulches, growing of live mulches, inter cropping and  cover crops.  Apart from the above allelopathy and mycoherbicides can play an important role in reducing the weed infestation.
All authors declare that they have no conflicts of interest.

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