Weather conditions in the study area
During the growing season of wheat (September-June), the average air temperature in the years of study was higher than the long-term average (2006/07-2015-16). Throughout the duration of the experiment, the highest total precipitation (419.40 mm) was recorded in the second growing season (2017-18) of wheat cultivation. Nevertheless, the precipitation in this year of study was not distributed evenly. The lowest precipitation level occurred in the 2016-17 season, when the total was lower by 176.49 mm than the long-term average for this period (Fig 1). According to
Rouabhi et al. (2019), Algeria will experience a decrease in precipitation and an increase in temperatures which will have a direct impact on agricultural productivity and food security.
Measurements variability
The analysis of variance showed a significant effect of tillage system, year and the interaction (tillage system × year) at (P<0.01) on the number of plants/ m² (NPM) and the number of tillers/ m² (NTM). The NMP and the NTM recorded under NT system with all the three crop rotation, were successively 28% and 78% higher than in CT. The difference between the two systems was more important during the drought season (2016-17). Under CT, the soil disturbance is more or less homogeneous manner which affects the seed bed quality and therefore the seeds are not placed in the same depth resulting in non-homogeneous emergence and low emergence density which can affect the yield. While under direct seeding system, the regularity of the seeding depth, the better placement of fertilizers and the good adherence of the seed to the soil
(Mekhlouf et al., 2011) increased significantly emergence rate.
The combined analysis of variance indicated significant effect of the tillage system and year on the plant height and beards length at (P<0.01). The 2017-18 cropping season was the most favorable environment to express the potential of plant height (62.73 cm) and the highest mean of length beards (10.99 cm) was expressed during the 2018-19 season. The longest plant and beard length mean value were recorded under CT system. The system tillage × year interaction was significant of plant height and non-significant of beards length.
The spike length was not significantly influenced by neither of the studied factors or their interaction. Nevertheless, the best mean value of spike length (6.11 cm) was obtained in 2018-19 cropping season and the spike length was higher under the NT system (6.15 cm). The same result obtained by
Wozniak et al. (2019) on the barley crop where he observed the longest spike of barley in direct seeding.
The combined analysis of variance indicated that tillage system had no significant effect on LCHC and LRWC. On the other hand, the year effect was significant at (P<0.001) on the two measured physiological traits. The interaction (tillage system × year) effect was significant (P<0.01) only on LCHC and its mean values averaged over years from 28.13 (SCMR) in 2016-17 cropping season to 21.53 SCMR in 2018-19 cropping season. The rotations W/W and W/T/P/W recorded higher LCHC under CT, while the rotations W/L/W had higher result of LCHC under NT system. High LCHC maintains wheat yields especially under drought and high temperature at the end of the cycle
(Shirvani et al., 2021). In our study, the LCHC mean under NT system (25.76 SCMR) was higher than the mean under the CT system (25.55 SCMR) indicating a positive effect of the NT system on wheat yield particularly under drought at the end of cycle, which is frequent in the study area.
Leaf relative water content that reflects the water status of the plant, averaged over year from 50.21% in the 2016-17 cropping season to 80.77% in 2018-19 cropping season. Leaf water content is closely to photosynthesis activity, as it can provide early warnings of drought
(Song et al., 2021). The long period of drought during the spring season 2016-17 had a significant impact on LRWC. On the other hand, the LRWC mean value under NT was higher than CT. This better hydratation of the plants under the NT system indicates also a lower impact of drought on the wheat culture which is one of the main advantages of the NT system. Regarding rotations, the rotation W/ L/ W and W/ T/ P/ W had a higher LRWC under CT but the rotation W/ W had higher result under NT system (Table 3). These results indicate that the expression of the physiological traits was affected by the environment and to the lesser extent by the tillage system.
The analysis of variance of number of spikes/m² (NSM) indicated a significant effect of tillage system and the season (P<0.001), while the effect of their interaction was not significant (Table 3). The NSM under NT system (262.31 spike/m²) was 20% higher comparatively to CT (221.31 spike/m²) which, is in accordance with the findings of
Benniou and Bahlouli (2015). As between the years, the NSM during 2019 was lower than the NSM during 2017 and 2018 (according to SNK test) which is mainly related to the important NPM during these years. This might be due to the most favorable climatic conditions, as well as the availability of nutrients (especially nitrogen) allowed to obtain the most important number of ears per m² (289.70) during this growing season (2017-18).
Seed yield was significantly influenced by the system tillage at (P<0.01) and the season at (P<0.001). In fact, the 2017-18 season, the wettest of three years under the study gave a much higher grain yield (2.56 t/ha) than the other two. The smaller differences (0.3 t/ha) in seed yield between 2018-19 and 2016-17 seasons can be ascribed to the rainfall distribution. Wheat grain yield following lens and pea exceeded the yield in the wheat monoculture. This result shows the importance of introducing legumes in a cropping system.
Kumar and Roy Sharma (2000) and
Sainju et al., (2017) proved that introducing legumes into the rotation can enrich the soil with N, break the cycle of diseases and weeds and conserve soil moisture which, resulted in a higher cereal yields after legumes. In our study, wheat grain yield under no-till system over the three rotations was 34% higher compared to CT. The presence of residues under NT system, increases soil moisture content and reduces water evaporation compared to CT system, especially during the dry season, can explain the increased wheat grain yield
(Lopez-Bellido et al., 1996).
Tillage system had no significant effect on 1000-seed weight; straw yield and biological yield, in the opposite of the year effect that was significant at (P<0.001) on these measures. Nevertheless, Straw yield and biological yield were higher under NT system. Seed weight was higher in W/W on NT system, but in the rotation W/L/W and W/T/P/W seed weight was higher under CT system. The biological yield and straw yield were higher in NT system on the rotations W/W and W/L/W but the best expression in W/T/P/W was obtained in CT (Table 2).
The harvest index (HI) was influenced significantly (P<0.001) by the tillage system, year and their interaction. This trait had a direct effect on cereal yield
(Kumar et al., 2021), it reflects the efficiency of photosynthesis products translocation to the grain and it severely affected by drought particularly at the end of cycle that characterizes the Mediterranean climate. Between the seasons, the highest HI was fetched in 2018-19 (0.33%) where the climate was favorable for production and the lowest was obtained in the season 2017-18 (0.13%). The HI recorded under zero tillage (0.27%) was significantly higher than HI under the CT (0.19 %). NT is known to improve water storage in soil which limits the impact of drought on production and the higher yields under NT system indicates a better water use efficiency (Table 2).
In addition, the seed yield was significantly and positively correlated with its components spike number per m² and 1000 seed weight and both yield and 1000 seed weight were also positively related with the plant height which could be considered as an indicator of favorable conditions for production (Table 4).